“I don’t think our audience is looking for unequivocal ‘rah-rah,’ ” said Ms. Maddow, who calls herself a liberal but not a Democrat.

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The spectacle of Democrats sniping at one another is not new, but having a TV home for it is. MSNBC — sometimes critically called the “home team” for supporters of Mr. Obama — has even hit upon the theme with a promotional tagline, “pushing back on the president,” in commercials for “Hardball,” Chris Matthews’s political hour.

“Our job is not to echo the president’s talking points,” said Phil Griffin, the president of MSNBC. “Our job is to hold whoever’s in power’s feet to the fire.”

But is it good business? MSNBC is projected to take in $365 million in revenue this year, roughly the same amount as last year, when the presidential election bolstered its bottom line. Three years ago, before making a left turn, it had revenue of about $270 million a year. MSNBC’s parent company, NBC Universal, is on the verge of being spun out of General Electric in a deal that would make Comcast its controlling entity.

Gary Carr, the executive director of national broadcast for the media buying agency TargetCast, said the opinions matter less than the ratings they earn. With cable’s prime-time opinion shows, “you’re reaching a lot of people,” he said.

It is certainly reaching the White House. Anita Dunn, the departing White House communications director, calls Mr. Olbermann and Ms. Maddow “progressive but not partisan,” and in doing so, distinguishes them from Fox News, which she considers a political opponent. The MSNBC hosts, she said in an e-mail message last month, “often take issues with the administration’s positions or tactics and are never shy about letting their viewers know when they disagree.”

Ms. Maddow said that apart from an off-the-record meeting between Mr. Obama and commentators that she attended last month, she has heard little from the White House.

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Mr. Griffin said, “We heard a whole lot more from the Bush White House.”

MSNBC’s liberal points of view have made the channel an occasional thorn in the side of G.E., but the channel has also fostered a diversity of opinions that people like Adam Green, the co-founder of the Progressive Campaign Change Committee, say were lacking in the past.

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“There’s been a huge market void for a long time,” Mr. Green said. Speaking of the MSNBC hosts, he said, “They are creating an environment where progressive thinkers and activists can thrive.”

Ms. Maddow, not surprisingly, agrees. “What looks like the middle of the country ought to look like the middle on TV,” she said in an interview this month.

She paused and added, “Maybe that would have helped us make better policy decisions in the country in the past.”

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Sitting down to a midnight dinner in the East Village after her program on a recent Thursday, Ms. Maddow had shed her suit for a T-shirt. Four minutes in, a fan asked for an autograph. “You’re doing great work,” he said while she signed her name.

MSNBC’s political tilt — and Ms. Maddow’s ascension to one of the most influential positions in progressive America — are still starkly new phenomena. A Rhodes scholar with liberal radio roots, Ms. Maddow started to host MSNBC’s 9 p.m. hour on the eve of last year’s presidential election, at a time when MSNBC was wrestling with its political identity. New viewers materialized overnight, peaking at nearly two million a night in October 2008. Without an election to drive viewership, her program averaged 880,000 viewers last month.

As her objections to the Stupak amendment (so named for Representative Bart Stupak, Democrat of Michigan) indicate, much of her work these days involves the Democratic health care overhaul. Ms. Maddow, Mr. Olbermann and Ed Schultz, the channel’s 6 p.m. host, formerly of Air America, have all exhorted Democrats to keep the public option.

Mr. Schultz started a broadcast last month by asking, “Where is the president? I think it’s time to be clear — crystal clear. What does Barack Obama want when it comes to health care in this country? What does he want in the bill?”

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Topics often tackled on Ms. Maddow’s program include the relationship between the United States military and politics (something she is writing a book about) and the repeal of the “Don’t ask, don’t tell” policy toward gays in the military.

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Representatives for two gay members of the military, Dan Choi and Victor Fehrenbach, approached Ms. Maddow’s producers about coming out on her show, in March and May respectively. Introducing Mr. Fehrenbach, Ms. Maddow intoned that he was about to be fired “in the shadow of these political promises left unfulfilled.”

Asked why she thought the two men had contacted her producers, Ms. Maddow said, “Maybe it’s because I’m gay; maybe it’s because we’ve covered this issue before on our air.”

Other MSNBC hosts have also objected to some of the president’s policy decisions. In April, Mr. Olbermann, the channel’s best-known voice, urged Mr. Obama to hold members of the Bush administration accountable for what he called the “torture of prisoners.”

“Prosecute, Mr. President,” he said. “Even if you get not one conviction, you will still have accomplished good for generations unborn.”

Ms. Maddow, however, contrasts her channel’s advocacy with the activism conducted, she says, by others on cable news. “We’re articulating liberal viewpoints,” she said at dinner, “but we’re not saying ‘Call your congressman, show up at this rally!’ ”

On her show, she has criticized Fox News for seeming to promote tea party rallies denouncing the administration this year, and the Fox host Glenn Beck, who has promoted a “9/12 Project,” intended, he says, to restore the values that Americans sensed immediately after the Sept. 11 terrorist attacks.

“I don’t have a constituency,” she said. “I’m not trying to form a — what would it be? — a ‘9/10’ movement.”